Revision to the Common Names of Butterflies 4

Revision to the Common Names of Butterflies
Part 4 : An Analysis of Name Changes

In Part 4 of our continuing discussion, analysis and recommendations of the butterflies' English Common Names, we complete the remaining species in the family Nymphalidae whose names have been amended by Dr Kirton. To reiterate our scope and biogeographic extent of the butterfly species, we had, established in our earlier discussions, the zoogeographical subregion of the Indo-Malayan ecozone, known as the Sundanian Subregion (or often called Sundaland) is the area of interest where species of butterflies have been assigned common names by various authors.




For the benefit of our readers who are viewing Part 4 of this discussion series, we would like to explain that a number of proposed changes were made by Dr Laurence Kirton in his recent book, A Naturalist's Guide to the Butterflies of Peninsular Malaysia, Singapore and Thailand. We analyse these changes and state our agreements or alternative views, and recommendations for future publications to consider.



Singapore species - Chocolate Sailor (Neptis harita harita)

We will now discuss the group of butterflies that come under the two genera Neptis and Phaedyma. If we refer to the earliest reference literature, The Identification of Indian Butterflies by Col W.H. Evans 1927, we note that the butterflies under these two genera comes under the collective generic name of "Sailers". Of interest to the Singapore butterfly fauna, are four species that fall under scrutiny.



Singapore species - Common Sailor (Neptis hylas papaja)

There has been quite a bit of debate as to the collective English common name of the Neptis and Phaedyma species. Certain countries refer to them as "Sailers", whilst others refer to them as "Sailors". In researching the basis of both names, I have come across arguments which support one or the other. It is interesting to note that the first option - Sailer was purportedly coined to describe the manner in which the butterfly flies i.e. it "sails".


A mating pair of Short Banded Sailors (Phaedyma columella singa)

This descriptor probably had some reference to the common names of some butterflies called "Gliders" also referring to the way the butterfly flies. In particular, searching for the name Neptis sappho would generally refer to a common name of Common Glider or Pallas Sailer. The name "Sailer" appears to be the preferred English common name used for the genera Neptis and Phaedyma in India, Sri Lanka, Bangladesh, Taiwan, Japan and Hong Kong.



The other English common name "Sailor" which refers to a seaman or mariner, is also often used as the collective common name for the Neptis and Phaedyma butterfly species. The reference to the name "sailor" could have originated from a number of sources. One origin could be the black-and-white striped wing patterns of the butterflies that are reminiscent of the colour of a navy sailor's uniform, which is white, with the navy blue collar detail.



The other possibility is related to the common English names that have their origins in military titles and ranks. In my earlier article on this blog, I made some postulations as to the origins of the butterflies' common names and their connections to British military titles and also gentry. Besides the land-based military titles, there are naval titles and naval ranks. After all, we have Admiral, Yeoman, Commodore and Commander - all of which are naval titles (mainly of the Royal Navy). So why not "sailors"?


A page from the book - Field Guide to the Butterflies of Southern Africa by Ivor Migdoll, Struik Publishers, Capetown, 1987 featuring Neptis saclava or the Small Spotted Sailor


A page from the book Common Butterflies of Vietnam by Alexander Monastyrskii and AlexeyDevyatkin, Labour and Social Affairs Publishing House, 2002.

The name "sailor" is the preferred English common name used by authors for Neptis and Phaedyma in Thailand, Vietnam, Malaysia, Borneo, Singapore, South Africa and China. In tracing the earliest reference book that uses the common name "sailor" for the Southeast Asian region, we have the "Butterflies of the Malay Peninsula" by Corbet and Pendlebury 1st Edition (1934). Hence there is undeniable evidence that even in the early 1930's or earlier, the British authors in Malaya who were writing their book were already using "Sailor" for these butterflies.


A page from Butterflies of the Malay Peninsula, 1st Edition by A S Corbet and H M Pendlebury, Kyle, Palmer and Co, 1934.

In his latest book, A Naturalist's Guide to the Butterflies of Peninsular Malaysia, Singapore and Thailand, Dr Kirton reverted to "Sailer" to describe the Neptis and Phaedyma species. Of all the authors of the references below which include Pisuth, Steven Neo, Yong Hoi-Sen, Ohtsuka, Morrell, Corbet and Pendlebury, Igarashi and Fukuda and many others covering the Southeast Asian region, the name "Sailor" is preferred.  Given that there are more books covering the Southeast Asian region using "Sailor" rather than "Sailer", we recommend to retain this common name for the Neptis and Phaedyma butterflies.

Recommendation : All the species of butterflies in the genera Neptis and Phaedyma to retain their collective English common name of "Sailors".


A page from Butterflies of the Malay Peninsula, 1st Edition by A S Corbet and H M Pendlebury, Kyle, Palmer and Co, 1934.

Amongst the Sailors in Singapore, there is one species, which warrants a change of name, if we take reference to the name given by the early publication for Southeast Asia, Butterflies of the Malay Peninsula by Corbet and Pendlebury, 1st Edition 1934. In the book, the species known then as Neptis nata cresina was given the common name Burmese Sailor.



Burmese Sailor (Neptis leucoporos cresina)

The scientific name of this species was later renamed to Neptis leucoporos cresina. It was, however called the Grey Sailor in Singapore. The recommendation should therefore be to refer to this species as Burmese Sailor as was the intention by the early authors who named Malayan butterflies back in 1934.

Recommendation : Neptis leucoporos cresina should be renamed Burmese Sailor.



Malay Baron (Euthalia monina monina)

The next butterfly name that was changed in Dr Kirton's book, is the Malay Baron. It was amended to Malayan Baron. The rationale cited for the change was "where the original name refers to a people group". In Part 3 of this series, we discussed the rationale for retaining "Malay" in the original common names of butterflies, instead of changing to "Malayan". Hence we recommend that the species Euthalia monina monina should retain its name Malay Baron, which is used in the majority of literature, papers and publications in the region.

Recommendation : Euthalia monina monina should retain its name Malay Baron.



Malay Viscount (Tanaecia pelea pelea)

The other related species that bears a similar background for a name change is the Malay Viscount. Hence by the same token mentioned above, the species Tanaecia pelea pelea should retain its well-known and widely-used name of Malay Viscount, instead of being changed to Malayan Viscount. Here, it should be clarified also that there are several other butterfly species bearing the name "Malayan". However in the case of the Malay Viscount, this is the name that it was referred to as early as the 1930's and we do not see a strong reason why it should be changed.

Recommendation : Tanaecia pelea pelea should retain its name Malay Viscount.



The next species of interest comes from the genus Lexias. Usually referred to as "Archdukes", these are medium-sized butterflies with robust bodies and are swift fliers. Dr Kirton amended the name of the species Lexias dirtea merguia from Black Tipped Archduke to Dark Archduke. In researching the rationale for this name change, we discovered an original name in Evans' book, Identification of Indian Butterflies, W.A. Evans, Diocesan Press, India, 1927, Dark Archduke, given to the species Adolias khasiana khasiana. The diagnostic description of the species corresponds with Lexias dirtea merguia (Black Archduke), and in particular where the description states that the "antennae black above".




In the same book, the name Archduke was given to the species with "antennae prominently yellow-tipped above" and described as Adolias dirtea jadeitina.  The two species were subsequently renamed from Adolias khasiana khasiana to Lexias dirtea merguia and Adolias dirtea jadeitina to Lexias pardalis dirteana.  The name Archduke will be retained for Lexias pardalis dirteana (previously Adolias dirtea jadetina) and the name Dark Archduke kept for Lexias dirtea merguia (previously Adolias khasiana khasiana).

Recommendation : Lexias dirtea merguia should revert to its original name of Dark Archduke.



The genus Eulaceura features only a single species in Malaysia and Singapore - Eulaceura osteria kumana. In Evans' book, the species is called the Elegant Emperor. This species was, however, called the Purple Duke in Singapore. Through regular usage, the Purple Duke name stuck. To be consistent with our policy of establishing the common name of the species based on the earliest valid publications for the Southeast Asian region, or, if unavailable, the reference to Evans' book, this species should henceforth revert to its original name of Elegant Emperor.



Eulaceura osteria kumana should now be called Elegant Emperor

It is also noted that Butterflies of Thailand, 2nd Edition by Pisuth Ek-Amnuay, also followed Evans' name for this species and called it Elegant Emperor. Further search did not turn up any other publications at this point in time that named this species as Purple Duke, except for Singapore-based publications and websites. It may take some time before the name is reinstated due to usage, but it would be appropriate to now call this species as Elegant Emperor.

Recommendation : Eulaceura osteria kumana should revert to its original English common name, Elegant Emperor.


Rounded Maplet (Chersonesia peraka peraka)

The next species in the series of renamed butterflies is the sole representative of the sub-family Cyrestinae in Singapore.  This species is Chersonesia peraka peraka. This species was given the name Little Maplet in Singapore references. Dr Kirton amended its common name to Rounded Maplet in his 2014 book. Upon checking with Evans, this name was in use since 1927. Butterflies of Thailand 2nd Edition also called this species Rounded Maplet.




Given that the English common name was originally known as Rounded Maplet, it would be appropriate to adopt this name henceforth to remain consistent with the relevant references in the region.

Recommendation : Chersonesia peraka peraka should revert to its original English common name, Rounded Maplet.



The final species of concern in the family Nymphalidae is one from the genus referred to collectively as Pansies. This species, Junonia hedonia ida, was not known from Evans time, as the species was absent from the Indian subcontinent. The closely related species, Junonia iphita, was given the common name Chocolate Soldier. There are some publications in the region which refers to Junonia iphita as Chocolate Pansy. However, since the earliest known coinage of the name for this species was Chocolate Soldier, that name should rightfully remain as such.



Chocolate Pansy (Junonia hedonia ida)

Hence the next most appropriate name for Junonia hedonia ida should be Chocolate Pansy. Dr Kirton amended the name of this species to Spotted Chocolate Soldier. Pisuth, in Butterflies of Thailand 2nd Edition, incorrectly named Junonia iphita as Chocolate Pansy. Perhaps it is because Junonia hedonia is unknown in Thailand. As Evans had precedence, then Junonia iphita should retain its name as Chocolate Soldier.  Hence Junonia hedonia should take the name Chocolate Pansy.

Recommendation : Junonia hedonia ida should retain its common name Chocolate Pansy.

This ends the common name changes in Dr Kirton's book for Nymphalidae. In the next article in the series, we will investigate the families Riodinidae and Lycaenidae.

Text by Khew SK : Photos by Sunny Chir, Goh EC, Khew SK, Loke PF, Nelson Ong, Horace Tan, Anthony Wong and Mark Wong

References :

[BPMST] A Naturalist's Guide to the Butterflies of P. Malaysia, Singapore & Thailand, Laurence G Kirton : John Beaufoy Publishing 2014
[C&P1] The Butterflies of The Malay Peninsula, A.S. Corbet and H.M. Pendlebury, 1st Edition, Kyle & Palmer, 1934.
[C&P4] The Butterflies of the Malay Peninsula, A.S. Corbet and H.M. Pendlebury, 4th Edition, Revised by Col John Eliot, Malaysian Nature Society, 1992
[BOT1] Butterflies of Thailand, Pisuth Ek-Amnuay, 1st Edition, Amarin Printing & Publishing, 2006
[BOT2] Butterflies of Thailand, Pisuth Ek-Amnuay, 1st Edition, Amarin Printing & Publishing, 2012
[CMB] Common Malayan Butterflies, R. Morrell, Longmans Malaysia, 1960
[MBAI] Malaysian Butterflies - An Introduction, Yong Hoi-Sen, Tropical Press, Malaysia, 1983
[BOS] Field Guide to the Butterflies of Singapore, Khew SK, Ink On Paper Publishing, Singapore, 2010
[BBSEA] Butterflies of Borneo & South East Asia, Kazuhisa Otsuka, Hornbill Books, Malaysia, 2001
[IIB] Identification of Indian Butterflies, W.A. Evans, Diocesan Press, India, 1927
[GCBOS] Guide to the Common Butterflies of Singapore, Steven Neo, Science Centre Singapore, 1996
[BSA] Butterflies of Southern Africa, Ivor Migdoll, Struik Publishers, Capetown, 1987
[BOV] Common Butterflies of Vietnam, Alexander Monastyrskii and AlexeyDevyatkin, Labour and Social Affairs Publishing House, 2002

Further Reading :

A Revision to the Common Names of Butterflies : Part 1
A Revision to the Common Names of Butterflies : Part 2
A Revision to the Common Names of Butterflies : Part 3

Taxonomic Classification in Butterflies

Taxonomic Classification in Butterflies
What is a Subspecies?

The Swedish naturalist, Carolus Linnaeus (Carl von Linne) pioneered the internationally recognised naming system for every creature and plant. Realising that every organism should be classified systematically according to its relationship with other organisms, he created the Linnaean system in his works, Systema Naturae in 1735 - a catalogue of the names of all known animals and plants. The 10th edition of Systema Naturae in 1758 is often adopted as the starting point for biological classification of animals and plants by family, genus and species.

According to the Linnaean system, all animals and plants are known by a combination of two names - the first indicating the genus, and the second, the species. This system, referred to as binominal nomenclature is also applied to the naming of butterflies. Furthermore, classification of butterflies used by many researchers and authors extend one more level to a rank below species. This is the subspecies. This method of naming the subspecies is known as the trinominal system and is now in general use for the scientific naming of butterflies.

A Pea Blue in Singapore is also known as a Long-Tailed Blue in Australia and Europe.  But its scientific name, Lampides boeticus, is consistent, wherever it is found.

I have sometimes been asked about what a subspecies is, and how a butterfly subspecies comes about. This blogpost discusses the definition of subspecies, and the way it is applied to the scientific naming of butterflies. Although most amateurs and hobbyists prefer to use English common names for butterflies, this can often be confusing, and common names tend to differ amongst different countries and published references. Scientific names, however, are adopted universally and Lampides boeticus is the name used for the species, whether in England or in Singapore. The English common name, however, may vary from Long-Tailed Blue to Pea Blue, depending on where the butterfly is sighted!

Before we examine the concept of subspecies, let's consider what makes a species. A species is a group of living things with very similar traits that can interbreed. For example, elephants can breed with each other, but not with hippopotamuses. Different species rarely interbreed. On the occasions when they do, either by human intervention or naturally, the offsprings of the interbreeding are typically sterile, and are unable to reproduce further.



By definition, a subspecies is an individual or a group within a species that has become somewhat physically different from the species. However, they are still biologically similar enough to interbreed with individuals of the same species. Now let's take a look at why some species branch into subspecies. One major reason for this is the geographic isolation of a particular population of that species e.g. by physical distance or geographical separation, like an island. They are still the same species, but the group that became isolated may take on some different physical features simply due to their location.


The southern Malaysian and Singapore subspecies malayana (left) has less dense striations on its wings than the northern Malaysian subspecies nina (right)

However, we need to understand that the science of taxonomy is ever-evolving. Scientists often find themselves in disputes about which genus a particular species belongs to, or whether two or more butterflies with very similar physical characteristics should be classified as separate species or as subspecies. With recent developments in microbiology, phylogenetics, DNA sequencing, and the study of the early stages of a butterfly often results in new insights on the relationships between species. This newfound knowledge regularly results in the necessity to make revisions to the adopted classifications of butterflies.

Scientific name of the Lime Butterfly is :
[Genus]Papilio [Species]demoleus [Subspecies]malayanus

In the trinominal system of naming of butterfly species, we have the [Genus] followed by the [species name] and then by the [subspecies name]. Hence for a Lime Butterfly, the scientific name would be Papilio demoleus malayanus. Note that only the genus name is capitalised. The species and subspecies name should never be capitalised.

Different subspecies of butterflies found in Singapore vs those found in Northern Thailand
Top : Moduza procris procis (Chiang Dao) vs Moduza procris milonia (Singapore)
Middle : Cheritra freja friggia (Singapore) vs Cheritra freja evansi (Chiang Dao)
Bottom : Lexias pardalis jadeitina (Chiang Dao) vs Lexias pardalis dirteana (Singapore)

For amateur observers (and even for the more knowledgeable enthusiasts), it is sometimes difficult to ascertain the physical differences between two subspecies of butterflies. Very often, purely due to geographical distances, a butterfly that shows very little perceptible physical differences is given a different subspecies name. Others, however, are more obvious where the physical differences are clear.

Examples of butterflies which has only the species name, suggesting that the physical appearance of these species have no significant differences irrespective of where they are found - Lampides boeticus (Pea Blue), Acreae terpsicore (Tawny Coster), Udaspes folus (Grass Demon) and Plastingia naga (Chequered Lancer)

Then what about butterflies with only species name given to it? This suggests that the species is consistent throughout its range and there are no discernible physical differences in its appearance, no matter where it is geographically found. Some examples of such butterflies that are extant in Singapore are the Pea Blue, Grass Demon, Tawny Coster and Chequered Lancer. Interestingly, quite a number of butterflies that have been described down to only species level are from the Lycaenidae and Hesperiidae families.

Left : Subspecies malayana and Right : subspecies parkeri are both found in Singapore

Left : Subspecies bolina and Right : subspecies jacintha are both found in Singapore

Left : Subspecies pratipa and Right : subspecies bisaltide are both found in Singapore. However, subspecies pratipa has not be reliably seen since 2007.

Left : Female subspecies bisalitide and Right : subspecies pratipa showing their differences on the upperside of their wings

In Singapore, there are 3 occurrences of butterflies of which there are two subspecies observed here over the years. These are the Knight (Lebadea martha parkeri and Lebadea martha malayana), the Great / Jacintha Eggfly (Hypolimnas bolina bolina and Hypolimnas bolina jacintha) and the Autumn Leaf (Doleschallia bisaltide bisaltide and Doleschallia bisaltide pratipa). Being an island separated from mainland Malaysia, it is likely that these subspecies' range overlap between Singapore and Malaysia and the evolution of the subspecies may continue until one is eventually eliminated.

Top : Caterpillar of the Autumn Leaf subspecies bisaltide and Bottom : subspecies pratipa

In the case of the Autumn Leaf, the subspecies pratipa which is usually found in Malaysia has a distinctively different caterpillar that is more obvious than the adult butterflies. However, subspecies pratipa was last seen in Aug 2007 at the Mandai Zoo and it is unknown if this subspecies still occurs in Singapore. The other two species continue to have sightings of their respective subspecies to this day.

As taxonomic science continues to evolve, technical methods and new knowledge are gained, it is inevitable that changes and updates to the scientific names of butterflies will continue.

Text by Khew SK : Photos by James Chia, Khew SK, Anthony Wong and Mark Wong