30 April 2026

Butterfly of the Month - April 2026

Butterfly of the Month - April 2026
The Orange Gull (Cepora iudith malaya)


We're coming to the end of one-third of the year 2026 and the world is still as volatile and turbulent as ever. Looking ahead to the remaining third of the year, will global peace return? Or we'll have some unexpected twists and turns in the world stability? Time will tell. As it is, time seems compressed as changes are the norm of the day and we have come to expect the unexpected as we chug along in our daily lives.

Orange Gulls puddling on sandy and muddy forest paths

Speaking of which, the US-Iran war continues to put a strain on oil prices and this is already hurting the pockets of the man-in-the-street thousands of kilometres away and who have nothing to do with the conflict. Prices of everyday goods and amenities continue to rise as the world economy depends heavily on the cost of fuel. The situation is reminiscent of the Oil Crisis of the 1970's, even as the world watches as the pendulum swings from war to peace and back to war in the Strait of Hormuz almost weekly.


Over in little Singapore, the pinch of our daily cost of living is growing. Prices of commodities and services are inching upwards slowly but surely. For the first time, the price of diesel exceeded the price of premium petrol at the local pumps. Traditionally, diesel was always seen as "inferior" fuel, pollutive and largely reserved for goods and industrial vehicles. Today, the price of diesel is higher than 98 octane petrol! This does not bode well, as diesel is the mainstay fuel for transportation, logistics and construction sectors and the knock-on effects will continue to be felt by the ordinary citizen in terms of cost of living.


Our feature butterfly for the month of April 2026 is the re-discovered Pierid, The Orange Gull (Cepora iudith malaya). Recorded by the early authors, this species was on the Singapore checklist based on past references. However, it has not been sighted reliably since the 1990's. However, in late 2018, the species was spotted on Pulau Ubin. Thereafter, it was regularly sighted on Pulau Ubin where a colony of the Orange Gull appeared to have established themselves. Thus far, it has not been spotted on Singapore island yet, and most sightings have been on Pulau Ubin.

Orange Gulls perched on the tops of leaves in the forest understorey when at rest

The Orange Gull is quite distinct in appearance where the tornal half of the upperside of the hindwing is a bright lemon yellow. The entire underside of the hindwing is yellow with brown veins and margins whilst the tornal area is a deeper orange-yellow. The three apical spots on the underside of the forewing are yellow.

Upperside of Orange Gulls - Top : Male, Bottom : Female

On the upperside, the male Orange Gull features a white forewing with prominent brown veins whilst the hindwing is white with the tornal area lemon yellow. The female is more brown dusted with the veins broader, whilst the hindwing has a brown marginal border and the lemon yellow tornal area is more reduced compared to the male.

Upperside of Orange Gulls - Top : Female, Bottom : Male

The species is described to be abundant in West Malaysia during the months of March and April and males can be observed puddling at sandy spots along forest streams and muddy footpaths. Females are rarer and found in the shaded forest understorey. Here in Singapore, the males are sometimes observed puddling along damp and muddy trails on Pulau Ubin. C&P5 describes the Orange Gull as "considered extirpated from Singapore". However, it is no longer the case, and the description should be updated when the book is revised in future.

The Bandicoot Berry (Leea indica) is a favourite nectaring plant of the Orange Gull on Pulau Ubin

Both sexes are regularly seen feeding on the flowers of the Bandicoot Berry (Leea indica) and sometimes perched on the tops of leaves with their wings folded upright when taking a rest in the shade. At certain hours of the day, both the males and females may be spotted sunbathing with partially opened wings. The butterfly is quick on the wing with an erratic flight, sometimes seen flying in the company of other Pierids like the Striped Albatross, Grass Yellows and Emigrants.


The life history of the Orange Gull has been successfully completed with the local host plants in Singapore recorded as Capparis micracantha (Capparaceae, common name: Thorn Caper, 小刺山柑), Crateva magna (Capparaceae, common name: Large Garlic Pear). The species has clearly established itself on Pulau Ubin as evident from the multiple observations of both adults and early stages at several sites on the island.

Text by Khew SK : Photos by Helen Chee, Choy CW, Ash Foo, David Ho, Khew SK, Low JK, Michael Khor, Michael Soh and Horace Tan

19 April 2026

Life History of the Ambon Onyx

Life History of the Ambon Onyx (Horaga syrinx maenala)


Butterfly Biodata:
Genus: Horaga Moore, 1881
Species: syrinx C. Felder, 1860
Subspecies: maenala Hewitson, 1869
Wingspan of Adult Butterfly: 23-26 mm
Caterpillar Local Host Plants: Melicope lunu-ankenda (Rutaceae), Clausena excavata (Rutaceae, common name: False Curry Leaf Tree), Ixonanthes icosandra (Ixonanthaceae, common name: Twenty-Man Tree), Citrus maxima (Rutaceae, common name: Pomelo), Citrus microcarpa (Rutaceae, common name: Calamansi), Ixora javanica (Rubiaceae, common name: Red Ixora).


A male Ambon Onyx on a leaf perch.

A female Ambon Onyx on a leaf perch.

A male Ambon Onyx perching on a small flower.

Physical Description of Adult Butterfly:
On the upperside, the male is bright clear blue and the female is purplish blue. The forewing has a broad black border and a narrow white discal patch crossed by dark-dusted veins. On the underside, both sexes are yellowish-brown with broad post-discal white bands traversing both fore- and hindwings. Both sexes have three filamentous tails on each hindwing, at veins 1b, 2 and 3, with the one at vein 2 longest among the three.

A female Ambon Onyx perching on a twig.

A male Ambon Onyx on a leaf perch.

Another male Ambon Onyx on a leaf perch.

Field Observations:
The Ambon Onyx is moderately common in Singapore, and sightings of the adults have been made in Southern Ridges, fringe of central catchment reserve and Pulau Ubin. It has a strong rapid flight, and is often observed to sunbathe at its favourite perches.

A male Ambon Onyx puddling on a mossy patch.

A male Ambon Onyx on a leaf perch.

A female Ambon Onyx perching on an Ixora flower.

Early Stages:
Caterpillars of Ambon Onyx is polyphagous and a number of larval host plants have been identified in Singapore. They are Melicope lunu-ankenda, Clausena excavata, Ixonanthes icosandra, Ixora javanica and Citrus spp. The caterpillars of the Ambon Onyx feed on the flowers, flower buds and young leaves of these host plants, with a strong preference for the flowers and flower buds.

Local host plant 1: Melicope lunu-ankenda.

Local host plant 2: Clausena excavata.

Local host plant 3: Ixora javanica.

A mating pair of the Ambon Onyx.

Eggs are laid singly on the young shoot or the inflorescence of the host plant. Each egg resembles a bun with coarse hexagonal reticulations. It is initially pale greenish when newly laid but turns whitish as it matures. Each egg has a basal diameter of about 0.8mm.

Two views of an egg of the Ambon Onyx.

A newly hatched caterpillar of the Ambon Onyx resting next to the empty egg shell.

The egg takes about 3 days to hatch. The caterpillar nibbles away the top part of the egg shell to emerge. The newly hatched is about 1.1mm long, with a pale yellowish brown head and a pale yellowish brown body covered with long setae dorsally and laterally. As growth progresses, dorsal tubercles on the metathorax (a pair) and 1st to 7th abdominal segments (one each) become marked in reddish brown. The first instar sees the body length reaches up to 3mm, and lasts about 2.5-3 days before the moult to the next instar.

Two views of a newly hatched caterpillar of the Ambon Onyx, length: 1.2mm.

Two views of a first instar caterpillar of the Ambon Onyx feeding on Red Ixora, length: 2.1mm.

Two views of a first instar caterpillar of the Ambon Onyx feeding on the young shoot of Twenty-Man Tree, length: 2.9mm.

Two views of a first instar caterpillar of the Ambon Onyx, length: 3mm.

Two views of a late first instar caterpillar of the Ambon Onyx, dormant prior to its moult.

The second instar caterpillar is either pale yellowish green or yellowish brown in base colour. It features numerous short setae on the body surface and has a number of prominent and pointed tubercles projecting from the body surface. The 2nd instar lasts about 2.5-3 days with the body length reaches up to 5-5.2mm.

Two views of an early second instar caterpillar of the Ambon Onyx feeding on Red Ixora, length: 3.3mm.

Two views of a second instar caterpillar of the Ambon Onyx feeding on Red Ixora, length:5mm.

Two views of a second instar caterpillar of the Ambon Onyx feeding on Twenty-Man Tree, length: 5mm.

Two views of a late second instar caterpillar of the Ambon Onyx, dormant prior to its moult.

The third instar caterpillar has the same configuration of dorsal, dorso-lateral and sub-spiracular tubercles as in the 2nd instar, but they are now proportionately longer and prominent, particularly so for the ones on 2nd, 6th and 7th abdominal segments. Whitish patches adorn the basal part of the dorsal tubercles. A whitish band also run sub-spiracularly along the side of the body. The third instar lasts about 3 days and has its length reaches up to about 10 mm before the moult to the 4th (and final) instar.

Two views of a third instar caterpillar of the Ambon Onyx feeding on Red Ixora, early in this instar, length: 5.6mm.

Two views of a third instar caterpillar of the Ambon Onyx, length: 9mm.

Two views of a late third instar caterpillar of the Ambon Onyx, dormant prior to its moult.

The 4th instar caterpillar has proportionately longer dorsal, dorso-lateral and sub-spiracular tubercles then those seen in the 3rd instar. Laterally, the lateral part of the 1st, 3rd, 4th and 5th abdominal segments are suffused with reddish brown to varying extent.

Two views of a 4th instar caterpillar of the Ambon Onyx, early in this instar, length: 10mm.

Two views of a 4th instar caterpillar of the Ambon Onyx, length: 11mm.

Two views of a 4th instar caterpillar of the Ambon Onyx, with much intense reddish coloration, length: 15mm.

A green form 4th instar caterpillar of the Ambon Onyx feeding on flowers buds of Melicope lunu-ankenda.

Two views of a 4th instar caterpillar of the Ambon Onyx, length: 16mm.

Two views of a 4th instar caterpillar of the Ambon Onyx feeding on a young leaf of the Twenty-Man Tree, green form, length: 18mm.

The 4th instar lasts about 4 days with the body length reaches up to 17-18mm. On the last day, the body gradually shortens. The caterpillar stops feeding, wanders around and eventually comes to rest on a spot on the stem or a pedicel where it begins the silk spinning effort to turn it into its pupation site. Once the silk pad is done, the caterpillar secures itself to it and turns into an immobile pre-pupa.

Two views of a pre-pupa of the Ambon Onyx, on a pedicel of Red Ixora.

The pre-pupal stage of the caterpillar lasts about one day. The ensuing pupation event turns it into a yellowish green pupa with the typical shape of a lycaenid pupa. The small pupa is about 8.5-9.5mm in length. The pupa attaches itself to the stem/pedicel via its broad cremaster end. The pupal surface is mostly smooth, with only a few short protrusions appearing on the dorsum in several abdominal segments. There is a pair of small reddish dorso-lateral patches on the 1st abdominal segment, and U-shaped pale brownish markings on the 3rd to 5th abdominal segments. Small irregularly shaped whitish patches also occur on the pupal surface and on the frontal part of the wing pads. In some pupae, the rear part of the wing pads is brownish. It is not uncommon for some pupae to feature little or no markings on the pupal surface.

Two views of a pupa of the Ambon Onyx, length: 9mm.

Two views of another pupa of the Ambon Onyx, length: 8.5mm.

Two views of another pupa of the Ambon Onyx, with reduced markings present.

After six to seven days, the pupa gradually turn dark towards the end of the day, with the wing pads prominently black with a few small whitish patches embedded. Next morning, the adult butterfly emerges from the mature pupa. It rests on the pupal case or nearby stem surface to have its wings gradually expanded. A few hours later, the adult Ambon Onyx takes its first flight.

Two views of a mature pupa of the Ambon Onyx.


A female Ambon Onyx emerging from its chrysalis.

A newly eclosed male Ambon Onyx resting near its pupal case.

A newly eclosed female Ambon Onyx resting near its pupal case.

References:
  • [C&P5] The Butterflies of The Malay Peninsula, A.S. Corbet and H.M. Pendlebury, G. and N. van der Poorten (Eds.), 5th Edition, Malayan Nature Society, 2020.
  • Butterflies of Thailand, Pisuth Ek-Amnuay, 2nd Edition, 2012.
  • A Field Guide to the Butterflies of Singapore, Khew S.K., Ink On Paper Communications, 2nd Edition, 2015.
Acknowledgment:
I would like to express my gratitude to Hanpeng Lim and Koh Cher Hern for sharing information on the host plant and early stages of the Ambon Onyx.

Text by Horace Tan, Photos by Koh Cher Hern, Loke PF, Nelson Ong, Anthony Wong, Khew SK and Horace Tan.