The Sea Almond (Terminalia catappa)
This 13th instalment of our Butterflies' Larval Host Plants series features Terminalia catappa
(Sea Almond), a member of Combretaceae which is a family of flowering plants comprising about 530 species of trees, shrubs and lianas. Sea Almond is a perennial and deciduous member which assumes the tree form.
The genus name "Terminalia" refers to the plants’ leaves being clustered at the end of branches.
The genus name "Terminalia" refers to the plants’ leaves being clustered at the end of branches.
Sea Almond is native to tropical Asia, North Australia and Polynesia. It has been introduced to and cultivated in many other tropical and sub-tropical regions. In Singapore, it can be found growing wild along the coast, on rocky and sandy beaches. It is also commonly planted in parks/gardens and along roadsides as ``shade'' trees.
Plant Biodata :
Family: Combretaceae
Genus: Terminalia
Species: catappa
Synonyms: T. procera, T. latifolia, T. moluccana.
Country/Region of Origin: Tropical Asia, North Australia and Polynesia.
English Common Name: Sea Almond
Other Common Names: Pacific Almond, Ketapang, Indian Almond, Tropical Almond Tree, 榄仁树.
Larval Host for Butterfly Species: Arhopala centaurus nakula (Centaur Oakblue), Flos apidanus saturatus (Plain Plushblue), Spindasis lohita senama (Long Banded Silverline).
Sea Almond is a fast growing tree with spreading canopy and a tiered growth form. It can grow to a height of 20-35m, and often has buttress roots. Its thin leathery leaves are large (8–38 cm long, 5–19 cm wide), spatula-shaped and spirally arranged at the end of the twigs. The leaves are shed up to twice a year with the colour turning yellow and read before dropping off.
Young developing leaves are yellowish brown to reddish, turning yellowish green to dark green when they mature. Leaves of Sea Almond contains tannins, and are used in tanning and dyeing. Other uses include treatment of headache and colic (using juice of young leaves), and topical treatment for rheumatic joints (in Indonesia and India).
Sea Almond is monoecious. Many tiny, white, petal-less flowers occur on long spikes emanating from leaf axils. Male flowers occur at the tip of the spike while the female flowers are at the bottom end.
Fruits are almond-shaped (3.5-7cm long), occurring in small clusters. Each fruit has a leathery, corky outer layer and a thick stony core containing the seed. It is initially green, turning yellow or red when ripened. The fruit is buoyant and can still viable after days in water.
The seed germinates with two near kidney-shaped cotyledons. Other young leaves follow thereafter.
In Singapore, the Sea Almond serves as the larval host plant for three butterfly species in the Lycaenidae family:
Centaur Oakblue,
Plain Plushblue ,
Long Banded Silverline.
Eggs of the three lycaenid species are laid singly or in small groups on young shoots, petioles or young leaves of the Sea Almond. Those of the Long Banded Silverline can also be found in crevices or cracks on the trunk.
Caterpillars of all three lycaenid species feed on young leaves of the Sea Almond. They live in close association with ants, and are typically sighted with a few ants in attendance. Caterpillars of the Long Banded Silverline even share residence with ants in latter's pavilion on the leaf underside. Caterpillars of the Centaur Oakblue and the Plain Plushblue typically build their own shelters on the leaf for rest and concealment. It is not unusual to find a fair number of ants in such leaf shelters.
References:
- The Concise Flora of Singapore : Hsuan Keng Singapore University Press, 1990.
- WildSingapore - Sea Almond.
- LKCNHM, The DNA of Singapore - Terminalia catappa.
- NPark's Flora Fauna Web - Terminalia catappa.
- The Plant Observatory - Terminalia catappa.
Text and Photos by Horace Tan.
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